Smithsonian Marine Station at Fort Pierce

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Themiste lageniformis

Photo of an adult sipunculan, Themiste lageniformis

Mary E. Rice

    Mary E. Rice
    Research Zoologist, Emeritus

  • Phone: (772) 462-0987
  • Fax: (772) 461-8154
  • E-mail:
  • Mailing and Shipping Address:
  • Dr. Mary E. Rice
    Smithsonian Marine Station
    701 Seaway Drive
    Fort Pierce, FL 34949 USA


University of Washington, Ph.D., 1966
Oberlin College, M.A., 1949
Drew University, B.A., 1947

Research Interests

Systematics and development of marine sipunculan worms
Development and larval biology of marine invertebrates, worldwide

Professional Activities

Member and past president (1979) of the American Society of Zoologists/Society for Integrative and Comparative Biology

Member and past president (1999) of The American Microscopical Society

Elected Fellow of the American Association for the Advancement of Science

Recent Publications

Boyle, Michael J. and Rice, Mary E. 2014. Sipuncula: an emerging model of spiralian development and evolution, The International Journal of Developmental Biology, 58:485-499

Young, Craig M., Emson, Roland H., Rice, Mary E. and Tyler, Paul A. 2013. A paradoxical mismatch of fecundity and recruitment in deep-sea opportunists: Cocculinid and pseudococculinid limpets colonizing vascular plant remains on the Bahamian Slope, Deep Sea Research Part II: Topical Studies in Oceanography, 92:36-45

Young, Craig M., He, Ruoying, Emlet, Richard B., Li, Yizhen, Qian, Hui, Arellano, Shawn M., Van Gaest, Ahna, Bennett, Kathleen C., Wolf, Maya, Smart, Tracey I. and Rice, Mary E. 2012. Dispersal of Deep-Sea Larvae from the Intra-American Seas: Simulations of Trajectories Using Ocean Models, Integrative and Comparative Biology, 52(4):483-496

Schulze, Anja, Maiorova, Anastassya, Timm, Laura E. and Rice, Mary E. 2012. Sipunculan Larvae and "Cosmopolitan" Species, Integrative and Comparative Biology, 52(4):497-510

Rice, Mary E., Reichardt, Hugh F., Piraino, Julianne and Young, Craig M. 2012. Reproduction, development, growth, and the length of larval life of Phascolosoma turnerae, a wood-dwelling deep-sea sipunculan, Invertebrate Biology, 131(3):204-215

Full Publications List

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