Smithsonian Marine Station at Fort Pierce

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Themiste lageniformis

Photo of an adult sipunculan, Themiste lageniformis

Michael J. Boyle

    Michael J. Boyle
    Research Biologist

  • Phone: (772) 462-0987
  • Fax: (772) 461-8154
  • E-mail:
  • Mailing and Shipping Address:
  • Dr. Michael J. Boyle
    Smithsonian Marine Station
    701 Seaway Drive
    Fort Pierce, FL 34949 USA


University of Hawaii at Manoa, Ph.D., 2010
Humboldt State University, B.Sc., 2003

Research Interests

Comparative morphological and molecular development of the Sipuncula
Developmental biology of marine invertebrate life histories
Embryology, evolution and diversity of larval forms

Professional Activities

Member of the Society for Integrative and Comparative Biology
Member of the American Microscopical Society

Recent Publications

Boyle, M.J., (Editor)and Kawauchi, G.Y. (contributing Editor): Proceedings of the 2nd International Symposium on the Biology of the Sipuncula. Smithsonian Institution Scholarly Press, Washington D.C. (in review).

Boyle, M.J., and Rice, M.E. Comparative Development and the Evolution of Life History Diversity in Sipuncula. In Proceedings of the 2nd International Symposium on the Biology of the Sipuncula. Smithsonian Institution Scholarly Press, Washington D.C. Editor: Boyle, M.J. (in review).

Lemer, S, Kawauchi, G.Y., Andrade, S.C.S., Gonzalez, V.L., Boyle, M.J., and Giribet, G. 2015. Re-evaluating the phylogeny of Sipuncula through transcriptomics. Molecular Phylogenetics and Evolution 83: 174-183.

Boyle, M.J., and Rice, M.E. 2014. Sipuncula: an emerging model of spiralian development and evolution. In Spiralian Model Systems Special issue of the International Journal of Developmental Biology 58: 485-499.

Boyle, M.J., Yamaguchi, E., and Seaver, E.C. 2014. Molecular conservation of metazoan gut formation: evidence from expression of endomesoderm genes in Capitella teleta (Annelida). EvoDevo 5: 39. Highly accessed

Herranz, M., Boyle, M.J., Pardos, F., and Neves, R.C. 2014. Comparative myoanatomy of Echinoderes (Kinorhyncha): a comprehensive investigation by CLSM and 3D reconstruction. Frontiers in Zoology 11: 31.

Hao, G., Boyle, M.J., Zhou, L., and Duan, Y. 2013. The intracellular citrus Huanglongbing Bacterium, Candidatus Liberibacter asiaticus encodes two novel autotransporters. PLoS ONE 8(7): e68921.

Herranz, M., Pardos, F., and Boyle, M.J. 2013. Comparative morphology of serotonergic-like immunoreactive elements in the Central Nervous System of kinorhynchs (Kinorhyncha, Cyclorhagida). Journal of Morphology 274 (3): 258-274.

Giani, V.C., Yamaguchi, E., Boyle, M.J., and Seaver, E.C. 2011. Somatic and germline expression of piwi during development and regeneration in the marine polychaete annelid Capitella teleta. EvoDevo 2: 10.

Meyer, N.P., Boyle, M.J., Martindale, M.Q., and Seaver, E.C. 2010. A comprehensive fate map by intracellular injection of identified blastomeres in the marine polychaete Capitella teleta. EvoDevo 1: 8.

Boyle, M.J., and Seaver, E.C. 2010. Expression of FoxA and GATA transcription factors correlates with regionalized gut development in two lophotrochozoan marine worms: Chaetopterus (Annelida) and Themiste lageniformis (Sipuncula). EvoDevo 1: 2.

Shimeld, S.M., Boyle, M.J., Brunet, T., Luke, G.N., and Seaver, E.C. 2010. Clustered Fox genes in lophotrochozoans and the evolution of the bilaterian Fox gene cluster. Developmental Biology 340: 234-248.

Boyle, M.J., and Seaver, E.C. 2009. Evidence of a dorsal pharynx in the marine polychaete, Capitella teleta (Polychaeta: Capitellidae). Zoosymposia 2: 317-328.

Boyle, M.J., and Seaver, E.C. 2008. Developmental expression of foxA and gata genes during gut formation in the polychaete annelid, Capitella sp. I. Evolution and Development 10: 89-105.

Boyle, M.J., Janiak, D., and Craig, S.F. 2007. Succession in a Humboldt Bay marine fouling community: the role of exotic species, larval settlement and winter storms. Proceedings of the 2004 Humboldt Bay Symposium. 215-233.

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